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Finding new species

By Jeremy Hance, mongabay.com
October 21, 2010


Species discovery: how do scientists find and describe new species—and the answer to other taxonomy questions

We are fortunate enough to share our world with an incredible array of wonderful and wild creatures. Yet despite centuries of scientists searching the world over for species and meticulously describing them, we still know surprisingly little about global biodiversity.

Currently scientists have described 1.9 million species from Balaenoptera musculus (the blue whale) to Oxysternon festivum (the red dung beetle), yet conservative estimates of the number of species on Earth range anywhere from 10 to 20 million species (some estimates jump to 100 million). If bacteria are added to this list then the number of species on Earth likely jumps another 10 million or so.


Gumprechts green pitviper (Trimeresurus gumprechti), which was described in 2002, is found across much of the Greater Mekong region. Image © Rene Ries.
In other words, at the very best—and not including bacteria—humans have identified only 20 percent of the species with which we share our planet. Whatever the percentage (and it's probably not nearly as high as 20), this means the discovery of new species will be on-going for some time now. In fact, if we take 2008 as an example—when researchers discovered just over 18,000 species—it would take 444 more years to uncover the world's additional 8 million (very conservative estimate) unknown species. If 50 million species inhabit the Earth, it would take researchers 2,500 more years to catalogue the world's species at current levels.

The search for species is even more pressing now that scientists believe we are either entering or already in the midst of a mass extinction. Due to a variety of human driven impacts—including deforestation, habitat loss, pollution, climate change, invasive species, and overexploitation such as through bushmeat or the pet trade—species populations are falling worldwide.


A new species of miniature frog was discovered in Borneo in 2010. Microhyla nepenthicola, shown here on the tip of a pencil, is about the size of a pea. © Indraneil Das/ Institute of Biodiversity and Environmental Conservation
The discovery of new species has also become important for many conservation organizations in attracting media attention and even funding. Few wildlife stories gain as much attention as new species and, it goes without saying, that certain new species (such as a new mammal) will receive more attention than others (a new mushroom).

But how do scientists find and describe new species?

The process of describing species

When describing a new species, scientists must collect a 'type specimen': this means a dead individual of the species. There have been a few instances where photographs including other genetic samples have been used to describe a new species successfully, and no individual is killed, but these are currently much in the minority.


Formally described in 2010, Durrell’s vontsira (Salanoia durrelli) was the first new carnivorous mammal to be discovered in Madagascar in 24 years. © Durrell Wildlife Conservation Trust.
The species must be closely examined to first determine that it doesn't in fact belong to another already-described species. For well-known species, such as a new mammal, this is easier than, for example, a rainforest insect. The specimen is compared to other similar species, including using descriptions and illustrations in published literature. If species appear very similar, genetic testing is increasingly used as a tool to determine new species.

Once the species is determined as new to science, researchers write a formal description of it, take photos and/or provide illustrations, and invent a new scientific name. After completing the paper, researchers then submit it to a scientific journal. The editor of the journal will circulate the new finding to other experts with those particular types of animals or plants. If the other experts agree that the new species is valid, the journal accepts the paper and the specimen officially becomes a new species. This process is long and often takes years between initial discovery and formal acceptance of the new species.

Cryptic species throw a wrench into this: these are species that look remarkably similar to another species, but are genetically unique. DNA is increasingly being used as a tool to determine new species as well as to discover the relationship between previously known species.

Finding species: where are the unknown species hiding?

If you want to earn scientific fame by discovering a new species, here are a few tips: think small, think underwater or underground, and think invertebrates and plants.

This is not to imply that all the world's large land-dwelling species have been found—just this year researchers announced the discoveries of a new species of monitor lizard that was as long as a adult human, a new monkey from the Brazilian Amazon, and a new carnivorous mammal from a lake in Madagascar—but such discoveries are more a matter of right place-right time over persistence.

While scientists still find new species in terrestrial ecosystems, a single dive to the depths of the ocean is likely to produce a new species. Cave systems have also yielded a number of new species, especially those that have only recently been explained. When it comes to types of species, insects are the most commonly discovered: thousands of new insects are found every year. Beetles are the most numerous.

History of species discovery and description

The history of species discovery and description is not only exciting—madness, exploration, untimely demise—it is also important for understanding the structure taxonomists use to describe species, for example the reason why every species is known by two Latinized names—the scientific name—one representing the genus and the other the species.

While the effort to categorize species stretches far back in history—Aristotle attempted it—today's modern system of categorizing and naming species largely stems from 18th Century Swedish scientist, Carl Linnaeus. In two seminal works—Species Plantarum and Systema Naturae—Linnaeus outlined the hierarchal structure employed today for classifying species. Latin was the language of choice, because it was dead and therefore fixed. Within his works, Linnaeus classified thousands of plants and animals from all around the world with Latinized names.

But Linnaeus didn't just build a structure to categorize species that would last to present-day, he was also instrumental in pushing the science of taxonomy further. As one of the most influential scientists of his day, Linnaeus chose and funded young biologist to travel the world, find new species, and return to him with specimens. These students have come to be known as 'the apostles of Linnaeus'. Seven of these 'apostles' died on their explorations, a couple went mad or became addicts, but many brought home species from as far away as China, South Africa, and the Amazon.

Linnaeus updated

While Linnaeus, with help from his traveling apostles, began the formal and large-scale effort to describe the world's life-forms, it has, of course, changed considerably in over two hundred years. When biologists refer to species today they often give the year of the species' formal description and the scientist who described it. For example, the well-known panda bear is Ailuropoda melanoleuca David, 1869.

Also contemporary scientists categorize a species according to where it fits in the evolutionary tree, a concept that would have been unknown to Linnaeus as he developed his system over a hundred years before Darwin outlined the theory of evolution. In other words, animals that share the same genus are more closely related in evolutionary terms than animals that share only the same family.

Instead of relying on five classifications, as Linnaeus did, researchers have developed a seven rung hierarchy to describe a unique life form: kingdom, phylum, class, order, family, genus, and species. For example, the tiger, which was described by Linnaeus, is classified in descending order as anamalia (animal), chordate (vertebrates), mammalia (mammals), carnivore (carnivores or mostly meat-eaters), felidae (cats), panthera (cats that roar), and tigris (the tiger). The tiger is additionally further separated into six surviving subspecies and three extinct subspecies.

Subspecies are a puzzling and at times difficult classification that establishes when a species has clearly splintered into distinct populations, usually geographically separated, but has not yet evolved far enough from each other to be considered separate species. One could think of a subspecies as unconnected populations: in other words the distinct populations would interbreed if they could, yet something—most often a physical barrier—is making interbreeding impossible. Debates over whether or not an animal is a unique species or simply a subspecies can be complex, rowdy, and linger for decades. This is because the debate over subspecies and species has real-world consequences: subspecies are rarely given conservation attention and funding. One example of a subspecies in flux is the forest elephant from the African Congo. The forest elephant is considered by some scientists to be distinct enough from the African elephant to be considered its own species, others, however, claim it is only a subspecies. If the forest elephant was accepted as a species today, the effort to save it from extinction would likely gain sudden life and an influx of funding. On the other side, if too many 'subspecies' are upgraded without cause, conservationists worry it could 'cheapen' the idea of a species.


Articles on recent species discoveries

Scientists discover world's deepest terrestrial animal
(02/22/2012) It's not the prehistoric monsters from the Jules Verne's Journey to the Center of the Earth or the human-bat hybrids of The Decsent, but it's an astonishing discovery nonetheless: intrepid scientists have discovered the world's deepest surviving terrestrial animal to date, a small wingless insect known as a springtail. Explorers discovered the new species, Plutomurus ortobalaganensis at a shocking 1.23 miles (1.98 kilometers) below the surface. The species was discovered by the Ibero-Russian CaveX Team Expedition in Krubera-Voronja Cave, the world's only known cave to go deeper than 2 kilometers.


Photo: new cookies-and-cream insect surprises researchers in Belize
(02/21/2012) Scientists have discovered the first ever insect in the Ripipterygidae family in Belize. Measuring only 5 millimeters (0.19 inches), the tiny insect uses its powerful legs to leap away from predators much like a grasshopper.


Photo: World's smallest chameleon discovered in Madagascar
(02/15/2012) Scientists have discovered four new species of super-tiny chameleons in Madagascar, according to a new paper in PLoS ONE. The smallest of the new species, Brookesia micra, is found only on the small island of Nosy Hara and has been dubbed the smallest chameleon in the world, measuring from nose to tail 29 millimeters (1.14 inches) at its largest. Scientists believe it represents a notable example of island dwarfism.


The camera trap revolution: how a simple device is shaping research and conservation worldwide
(02/14/2012) I must confess to a recent addiction: camera trap photos. When the Smithsonian released 202,000 camera trap photos to the public online, I couldn’t help but spend hours transfixed by the private world of animals. There was the golden snub-monkey (Rhinopithecus roxellana), with its unmistakably blue face staring straight at you, captured on a trail in the mountains of China. Or a southern tamandua (Tamandua tetradactyla), a tree anteater that resembles a living Muppet, poking its nose in the leaf litter as sunlight plays on its head in the Peruvian Amazon. Or the dim body of a spotted hyena (Crocuta crocuta) led by jewel-like eyes in the Tanzanian night. Or the less exotic red fox (Vulpes vulpes) which admittedly appears much more exotic when shot in China in the midst of a snowstorm. Even the giant panda (Ailuropoda melanoleuca), an animal I too often connect with cartoons and stuffed animals, looks wholly real and wild when captured by camera trap: no longer a symbol or even a pudgy bear at the zoo, but a true animal with its own inner, mysterious life.


Photo: new blue, red, yellow lizard discovered in the Andes
(02/13/2012) Researchers have discovered a new species of lizard in the Peruvian Andes, whose males sport beautiful colors, according to a paper in ZooKeys. The highest-dwelling known species of the genus Potamites, the new lizard, dubbed Potamites montanicola, was found in forest streams at 1,500 to 2,000 meters (4,900 to 6,500 feet). The species was discovered as apart of a biodiversity monitoring program by COGA, a Peruvian fossil fuel company.


Vampire and bird frogs: discovering new amphibians in Southeast Asia's threatened forests
(02/06/2012) In 2009 researchers discovered 19,232 species new to science, most of these were plants and insects, but 148 were amphibians. Even as amphibians face unprecedented challenges—habitat loss, pollution, overharvesting, climate change, and a lethal disease called chytridiomycosis that has pushed a number of species to extinction—new amphibians are still being uncovered at surprising rates. One of the major hotspots for finding new amphibians is the dwindling tropical forests of Southeast Asia.


Photos: 46 new species found in little-explored Amazonian nation
(01/25/2012) South America's tiniest independent nation still hides a number of big surprises: a three week survey to the sourthern rainforests of Suriname found 46 potentially new species and recorded nearly 1,300 species in all. Undertaken by Conservation International's (CI) Rapid Assessment Program (RAP) the survey found new species of freshwater fish, insects, and a new frog dubbed the "cowboy frog" for the spur on its heel. While Suriname may be small, much of its forest, in the Guyana Shield region of the Amazon, remains intact and pristine. The Food and Agriculture Organization (FAO) estimates that 91 percent of Suriname is covered in primary forests, however this data has not been updated in over two decades.


Scientists discover over 19,000 new species in 2009
(01/19/2012) In 2009 researchers described and named 19,232 species new to science, pushing the number of known species on Earth to just under two million (1,941,939 species), according to the State of Observed Species (SOS). Discoveries included seven new birds, 41 mammals, 120 reptiles, 148 amphibians, 314 fish, 626 crustaceans, and 9,738 insects.


New book series hopes to inspire research in world's 'hottest biodiversity hotspot'
(01/17/2012) Entomologist Dmitry Telnov hopes his new pet project will inspire and disseminate research about one of the world's last unexplored biogeographical regions: Wallacea and New Guinea. Incredibly rich in biodiversity and still full of unknown species, the region, also known as the Indo-Australian transition, spans many of the tropical islands of the Pacific, including Indonesia's Sulawesi, Komodo and Flores, as well as East Timor—the historically famous "spice islands" of the Moluccan Archipelago—the Solomon Islands, and, of course, New Guinea. Telnov has begun a new book series, entitled Biodiversity, Biogeography and Nature Conservation in Wallacea and New Guinea, that aims to compile and highlight new research in the region, focusing both on biology and conservation. The first volume, currently available, also includes the description of 150 new species.


Photos: program devoted to world's strangest, most neglected animals celebrates five years
(01/16/2012) What do Attenborough's echidna, the bumblebee bat, and the purple frog have in common? They have all received conservation attention from a unique program by the Zoological Society of London (ZSL) called EDGE. Five years old this week, the program focuses on the world's most unique and imperiled animal species or, as they put it, the most Evolutionarily Distinct and Globally Endangered (EDGE) species. In the past five years the program has achieved notable successes from confirming the existence of long unseen species (Attenborough's echidna) to taking the first photos and video of a number of targeted animals (the purple frog).


Beyoncé honored with new horse fly named after her
(01/16/2012) Musical artists, and dancer extraordinare, Beyoncé has been awarded a new honor this week: entomologists in Australia have named a new horse fly after the American singer. The new horse fly, dubbed Scaptia beyonceae, is found in Queensland's Atherton Tablelands.


Camera traps snap first ever photo of Myanmar snub-nosed monkey
(01/10/2012) In 2010 researchers described a new species of primate that reportedly sneezes when it rains. Unfortunately, the new species was only known from a carcass killed by a local hunter. Now, however, remote camera traps have taken the first ever photo of the elusive, and likely very rare, Myanmar snub-nosed monkey (Rhinopithecus strykeri), known to locals as mey nwoah, or 'monkey with an upturned face'. Locals say the monkeys are easy to locate when it rains, because the rain catches on their upturned noses causing them to sneeze.


Photos: scientists find new species at world's deepest undersea vent
(01/10/2012) It sounds like a medieval vision of hell: in pitch darkness, amid blazing heat, rise spewing volcanic vents. But there are no demons and devils down here, instead the deep sea vent, located in the very non-hellish Caribbean sea, is home to a new species of pale shrimp. At 3.1 miles below (5 kilometers) the sea's surface, the Beebe Vent Field south of the Cayman islands, is the deepest yet discovered.


Photo: Tiny lemur discovered in Madagascar forest
(01/08/2012) A new species of mouse lemur has been discovered in eastern Madagascar, report researchers from Germany. The species is described in a recent issue of the journal Primates.


'Lost world' dominated by Yeti crabs discovered in the Antarctic deep
(01/03/2012) Scientists have discovered a deep sea ecosystem dominated by hairy pale crabs off of Antarctica. The new species of "Yeti crabs" survive alongside many other likely new species, including a seven-armed meat-eating starfish, off of hydrothermal vents, which spew heat and chemicals into the lightless, frigid waters. According to the paper published in PLoS ONE, this is the first discovery of a hydrothermal vent ecosystem in the Southern Ocean though many others have been recorded in warmer waters worldwide.


The biggest new species discoveries in 2011
(12/26/2011) Every year scientists describe thousands of species for the first time. 2011 was no different, so here's a look at some of the significant new species discoveries for the year. New species discoveries are bound to continue as genetic analysis becomes more widespread and scientists conduct surveys of ever more habitats. But species loss also continues, and that is something considerably more difficult to quantify. Cataloging the disappearance of a species is extremely costly -- final proof can take years. Nonetheless in 2011 the book was closed on two particularly conspicuous creatures: neither the Vietnamese rhino nor the western black rhino roam the wilds any more.


The world's tiniest frogs, the size of a Tic Tac, discovered in New Guinea
(12/16/2011) Scientists have discovered the world's tiniest frogs in Papua New Guinea.


New large horned viper discovered, but biologists keep location quiet
(12/15/2011) In a remote forest fragment in Tanzania, scientists have made a remarkable discovery: a uniquely-colored horned viper extending over two feet long (643 millimeters) that evolved from its closest relative over two million years ago. Unfortunately, however, the new species—named Matilda's horned viper (Atheris matildae)—survives in a small degraded habitat and is believed to be Critically Endangered. Given its scarcity, its discoverers are working to pre-empt an insidious threat to new species.


Photos: 208 species discovered in endangered Mekong region in 2010
(12/14/2011) Last year researchers scoured forests, rivers, wetlands, and islands in the vanishing ecosystems of the Mekong Delta to uncover an astounding 208 new species over a twelve month period. A new report by the World Wide Fund for Nature (WWF) highlights a number of the new species—from a new snub-nosed monkey to five new meat-eating pitcher plants to a an all-female, cloning lizard—while warning that many of them may soon be gone as the Mekong Delta suffers widespread deforestation, over-hunting and poaching, massive development projects, the destruction of mangroves, pollution, climate change, and a growing population.


New species of frog sings like a bird
(12/12/2011) If you're trudging through the high-altitude rainforests of northern Vietnam and you hear bird song, you might want to check the trees for frogs. Yes, that's right: frogs. A new species of tree frog has been discovered in Vietnam that researchers say has a uniquely complex call that makes it sound more like a bird than a typical frog. Discovered in Pu Hoat Proposed Nature Reserve, the new species, dubbed Quang's tree frog (Gracixalus quangi), dwells in the forests at an altitude 600-1,300 meters (nearly 2,000-4,265 feet).


Photos: two new paper clip-sized frogs discovered in Vietnamese mountains
(12/07/2011) Researchers have discovered two new frog species living in the montane tropical forests of Vietnam. Known as moss frogs, these small amphibians employ camouflage as one way to keep predators at bay, in some cases resembling the moss that gives them their name.


Yeti crab cultivates bacteria on its claws to feed itself
(12/02/2011) A species of deep-sea crab found in hydrothermal vents off Costa Rica cultivates "gardens" of bacteria on its claws to feed itself, reports Nature News.


Photos: bizarre new mammal discovered in Vietnam
(11/28/2011) A new species of omnivorous mammal has been found in Cuc Phuong National Park, Vietnam. A part of the weasel family, the new species is known as a ferret-badger, a strange subfamily of mammals that have been little-studied by researchers. Smaller than proper badgers, ferret-badgers have elongated faces and long bushy tails.


Photos: two dozen new beetles discovered in Papua New Guinea hotspot
(11/23/2011) Over the past two decades, at least 24 new beetles species have been discovered in a remote mountainous rainforest region of Papua New Guinea by Swedish entomologists Ulf Nylander. Described in the new book Biodiversity, Biogeography and Nature Conservation in Wallacea and New Guinea, the new beetles found in the Aseki Province are all ecologically linked to rotting wood.


One night only: new orchid species surprises scientists
(11/22/2011) A mysterious new orchid blooms for one night only, opening around 10 PM and closing at 10 AM. Discovered on the island of New Britain near Papua New Guinea, the new species is the world's first orchid that flowers only at night. Scientists found the new flower, named Bulbophyllum nocturnum, in a logging concession on the tropical island.


Photos: bizarre shell of new snail baffles researchers
(11/10/2011) A new species of snail with a bizarre shell has surprised scientists. Discovered near massive waterfalls in pristine lowland rainforest in New Guinea, the tiny new species' shell is shaped like a cornucopia, spirals flying freely instead of fused together like most shells. Latvian malacologist (one who study molluscs) Kristine Greke, who described the new species, named it Ditropopsis mirabilis, meaning miraculous or extraordinary. To date, scientists are uncertain why the super small snail—2 to 6 millimeters (0.07 to 0.23 inches)—would have evolved such a strange shell.


Beetle bonanza: 84 new species prove richness of Indo-Australian islands
(11/08/2011) Re-examining beetle specimens from 19 museums has led to the discovery of 84 new beetle species in the Macratria genus. The new species span the islands of Indonesia, New Guinea, and the Solomon Islands, tripling the number of known Macratria beetles in the region. "Species of the genus Macratria are cosmopolitan, with the highest species diversity in the tropical rainforests. Only 28 species of this genus were previously known from the territory of the Indo-Australian transition," Dr. Dmitry Telnov with the Entomological Society of Latvia, who discovered the new species, told mongabay.com.


Photos: three bizarre bats discovered in Southeast Asia
(10/30/2011) In the forests of Cambodia and Vietnam, researchers have discovered three new species of tube-nosed bats, known for extraordinary nostrils that look like blooming flowers. The new bats, described in the Journal of Mammalogy, are likely imperiled by deforestation. "They all possess specially shaped nostrils (hence the name for the group) the exact role of which not known yet," Gabor Csorba, lead author of the paper with the Hungarian Natural History Museum, told mongabay.com.


Giant one-celled organisms discovered over six miles below the ocean's surface
(10/23/2011) Imagine a one-celled organism the size of a mango. It's not science fiction, but fact: scientists have cataloged dozens of giant one-celled creatures, around 4 inches (10 centimeters), in the deep abysses of the world's oceans. But recent exploration of the Mariana Trench has uncovered the deepest record yet of the one-celled behemoths, known as xenophyophores.


If camera traps don't prove existence of Bigfoot or Yeti nothing will
(10/13/2011) Let me state for the record that I am skeptical of the existence of Bigfoot or the Yeti, however I do have a fascination for following the latest news on the seemingly never-ending search for these hidden hominids. This week a Yeti conference in Russia announced 'indisputable proof' of the legendary hairy ape in the wilds of Southern Siberia. What did this proof consist of? Not DNA, photographs, video, or the Yeti itself (dead or alive) as one would expect from the word 'indisputable', but a few alleged Yeti hairs, an alleged bed, and alleged footprints. Cryptozoologists, those who are fascinated by hidden species such as the proposed Yeti and Bigfoot, don't serve their cause by stating the reality of a species without the evidence long-deemed necessary by scientific community to prove it—either a body or DNA samples combined with clear photographic evidence—instead they make themselves easy targets of scorn and ridicule.


Scientists confirm ancient Egyptian knowledge: Nile crocodile is two species
(09/20/2011) DNA has shown that the Nile crocodile is in fact two very different species: a bigger, more aggressive crocodile and a smaller, tamer species that today survives only in West Africa. While the taxonomy of the Nile crocodile has been controversial for over a century, the new study points out that the ancient Egyptians recognized the differences in the species and avoided the big crocodile for its rituals.


Pictures: 12 new species of frog discovered in India
(09/15/2011) Scientists have discovered 12 new species of frogs in the rainforests of India's Western Ghats, according to a paper published in the latest issue of ZooTaxa.


New species of bottlenose discovered in Australia (PHOTO)
(09/15/2011) Researchers have discovered a new species of dolphin in Australia, reports ABC News.


New 'demon' bat discovered in Vietnam (PHOTO)
(09/01/2011) Scientists have discovered three previously unknown bat species in southern Indochina, reports Fauna & Flora International.


New seabird discovered from Hawaii, but no one knows where it lives
(08/30/2011) Researchers have uncovered a new seabird native to Hawaii stuffed in a museum. Originally identified as a smaller variation of a little shearwater (Puffinus assimilis), DNA tests showed that the bird, which was collected over four decades ago, was in fact a unique species. Named Bryan’s shearwater (Puffinus bryani), the fate of this bird in the wild remains unknown.


Meet the just discovered 'Komodo dragon' of wasps
(08/28/2011) A new species of warrior wasp has been discovered on the Indonesian island of Sulawesi that is so large and, frankly, terrifying-looking that it has been dubbed the 'Komodo dragon' of the wasp family. Bizarrely, the male of the species has jaws that outstretch its limbs. "I don't know how it can walk," said the wasp's discoverer, entomologist Lynn Kimsey of the University of California, Davis and director of the Bohart Museum of Entomology, in a press release. "Its jaws are so large that they wrap up either side of the head when closed."


Photo: new titi monkey discovered in Amazon area under siege
(08/25/2011) A new species of titi monkey has been discovered in the Brazilian Amazon. Found during a 2010 December expedition, this is the second new titi monkey discovered in the Amazon in three years. In 2008 another new titi, dubbed the Caquetá titi, was discovered in the Colombian Amazon, although it was only announced last year. An expedition backed by WWF-Brazil found the new titi between the Guariba River and the Roosevelt River in northwestern part of Mato Grosso, a state of Brazil known as a center of Amazon destruction.


Humanity knows less than 15 percent of the world's species
(08/23/2011) Scientists have named, cataloged, and described less than 2 million species in the past two and a half centuries, yet, according to an new innovative analysis, we are no-where near even a basic understanding of the diversity of life on this small blue planet. The study in PLoS Biology, which is likely to be controversial, predicts that there are 8.7 million species in the world, though the number could be as low as 7.4 or as high as 10 million. The research implies that about 86 percent of the world's species have still yet to be described.


New species is eel-equivalent of the coelacanth
(08/18/2011) The ocean holds endless surprises still. In an underwater cave off the Pacific island nation of Palau, reachers have made an astounding discovery: an eel species unknown to science that harkens back 200 million years. The species, described in the Proceedings of the Royal Society B as an 'enigmatic, small eel-like fish', sports anatomical features that differentiate it from the over 800 known species of eel surviving today.


Featured video: WWF's Astonish Me
(08/16/2011) Highlighting new species recently discovered around the world, the short film Astonish Me, was created as apart of a happy 50th birthday celebration for conservation organization WWF.


Photos: 10 new frogs discovered in India's great rainforest
(08/09/2011) Ten new species of frog have been discovered in India's Western Ghats according to two new papers in Biosystematica. Although human populations have farmed in the Western Ghats for centuries, the new discoveries prove that the rainforest still holds many surprises. The Western Ghats lie along India's west coast and have been dubbed one of the world's biodiversity hotspots, but the rich wildlife is imperiled by rising human impacts.


Photo: six new mini-moths discovered
(07/19/2011) Researchers have discovered six new species of moth from Central America, according to a new paper in Zoo Keys. The moths belong to the primitive Yponomeutidae family, which are commonly known as ermine moths, since some of the species' markings resemble the coat of the ermine.


Photo: four new jewel beetles uncovered in Thailand and Indonesia
(07/10/2011) Researchers have discovered four new species of jewel beetles, one from Thailand and three from Indonesia. Jewel beetles, in the beetle family Bupretidae, are known for their iridescent colors.


Photo: New mouse species discovered in Brazil
(07/06/2011) Researchers discovered a new species of mouse in Brazil, reports the Chico Mendes Institute for Biodiversity Conservation (ICMBio).


2-4 new shrew species discovered in Sulawesi
(06/28/2011) A research expedition has turned up two to four new species of shrew on the Indonesian island of Sulawesi, reports a conservation group working to protect their forest habitat.


Pictures: Turquoise 'dragon' among 1,000 new species discovered in New Guinea
(06/27/2011) Scientists discovered more than 1,000 previously unknown species during a decade of research in New Guinea, says a new report from WWF. While the majority of 1,060 species listed are plants and insects, the inventory includes 134 amphibians, 71 fish, 43 reptiles, 12 mammals, and 2 birds. Among the most notable finds: a woolly giant rat, an endemic subspecies of the silky cuscus, a snub-fin dolphin, a turquoise and black 'dragon' or monitor lizard, and an 8-foot (2.5-m) river shark.


Photos: 300 species discovered during expedition to Philippines
(06/26/2011) Scientists believe they have discovered more than new 300 species during a six-week expedition to the Philippines.


7 new mice species discovered in the Philippines
(06/16/2011) Seven new species of mice have been discovered in the rainforests of Luzon island in the Philippines, according to the country's Department of Environment and Natural Resources.


Photo: Scientists discover 'SpongeBob' mushroom in Borneo
(06/16/2011) Scientists have discovered a colorful new species of mushroom in the rainforest of Borneo and named it after a popular cartoon character: SpongeBob.


New bee species sports world's longest tongue
(06/14/2011) A new species of bee discovered in the Colombian rainforest could give the world's biggest raspberry! Researchers say the new bee has the longest tongue of any known bee, and may even have the world's longest tongue compared to body size of any animal: twice the length of the bee itself. The new species has been named Euglossa natesi in honor of bee-expert Guiomar Nates.


Photos: 600 new species discovered in Madagascar since 1999
(06/06/2011) More than 600 species of plants and animals have been described in Madagascar over the past decade, reiterating the position of Indian Ocean island as one of the world's top biodiversity hotspots, says a new report issued today by WWF.


Photos: new bat uncovered in the Caribbean
(05/26/2011) Researchers have declared a new species of bat from the Caribbean island of St. Vincent. While the new bat had been documented before, it was long believed to be a member of a similar species that is found throughout South America and a few Caribbean Islands, that is until PhD student Peter Larsen noticed it was far larger than its relative down south.


Photos: the top ten new species discovered in 2010
(05/23/2011) If we had to characterize our understanding of life on Earth as either ignorant or knowledgeable, the former would be most correct. In 250 years of rigorous taxonomic work researchers have cataloged nearly two million species, however scientists estimate the total number of species on Earth is at least five million and perhaps up to a hundred million. This means every year thousands of new species are discovered by researchers, and from these thousands, the International Institute for Species Exploration at Arizona State University selects ten especially notable new species.


3,000 amphibians, 160 land mammals remain undiscovered—that is if they don't go extinct first
(05/18/2011) Remote little-explored rainforests probably harbor the majority of undiscovered amphibians and land mammals according to a new study in Proceedings of the Royal Society B. The study calculated that it's likely 33% of the world's amphibians and 3% of terrestrial mammals still remain unknown. However, the paper also found that these undiscovered species are likely in worse peril of extinction than already-described species.


No limbs or sight needed: bizarre new lizard uncovered in Cambodia
(05/09/2011) A new species of legless lizard has been discovered in Cambodia. Herpetologist Neang Thy uncovered, literally, the new species when he turned over a log in the species-rich Cardamom Mountains. While the new lizard looks like a snake or a big earthworm, it is in fact a lizard belong to the Dibamidae family. These bizarre reptiles spend much of their lives burrowing underground for insects, which has allowed them to lose the need for limbs.


A new rhino species?
(04/11/2011) Using genetic data and re-assessing physical evidence, scientists write that they have uncovered a new species of rhino, long considered by biologists as merely a subspecies. Researchers write in an open access PLoS ONE paper published last year that evidence has shown the northern white rhino is in fact a distinct species from the more commonly known—and far more common—southern white rhino. If the scientific community accepts the paper's argument it could impact northern white rhino conservation, as the species would overnight become the world's most endangered rhino species with likely less than ten surviving.


The saola: rushing to save the most 'spectacular zoological discovery' of the 20th Century
(04/04/2011) The saola (Pseudoryx nghetinhensis) may be the most enigmatic, beautiful, and endangered big mammal in the world—that no one has ever heard of. The shy ungulate looks like an African antelope—perhaps inhabiting the wide deserts of the Sahara—but instead it lives in the dense jungles of Vietnam and Laos, and is more related to wild cattle than Africa's antelopes. The saola is so unusual that is has been given its own genus: Pseudoryx, due to its superficial similarities to Africa's oryx. In the company of humans this quiet forest dweller acts calm and tame, but has yet to survive captivity long. Yet strangest of all, the 200 pound (90 kilogram) animal remained wholly unknown to science until 1992.


'Luck and perseverance': new plant genus discovered in Amazon
(03/31/2011) The discovery of a new plant species is not uncommon, especially in places of remarkable biodiversity such as the Amazon rainforest. However, discovering a new plant genus, a taxonomic rank above species, is, according to Henk van der Werff fromt the Missouri Botanical Garden (MBG), "a matter of luck and perseverance". Researchers with the Missouri Botanical Garden have been blessed with both as they have announced two new species of Amazonian plants, one from Ecuador and one from Peru, that comprise a completely new genus: named, Yasunia, since the plant was originally discovered in Ecuador's vast Yasuni National Park.


Photo: new vipers discovered in Asia's rainforests
(03/30/2011) Researchers have discovered two new species of pitviper in Southeast Asia. After collecting snakes throughout the Asian tropics—Thailand, Laos, Vietnam, and Cambodia—researchers were able to parse out a more complex set of species than had been recognized. One of the new vipers has been dubbed Cryptelytrops rubeus for its ruby-colored eyes.


New seabird discovered, first in 55 years
(03/23/2011) Stephen Maturin, if he were not fictional, would be delighted. A new seabird has been discovered by an international expedition headed by one of the world's top seabird-experts, Peter Harrison, after he received photos from vacationing birders of an unusual looking storm petrel off the coast of Chile.


Photos: two new freshwater stingrays discovered in the Amazon
(03/09/2011) Few people probably realize that in the rivers and lakes of the Amazon rainforest large stingrays glide, searching for crustaceans and small fish. Equipped with a powerful barbed tail they are often feared by locals. However, even as big as these fish are, new species continue to be described. Recently, scientists have identified two new species of Amazonian freshwater stingray near Iquitos, Peru. The new stingrays are unique enough to be placed in a new genus (the taxonomic level above species) called Heliotrygon, the first new Amazonian stingray genus to be described in nearly 25 years.


New species of zombie-creating fungi discovered
(03/02/2011) As everyone knows, human zombies are created when an uninfected human is bitten by a member of the brain-craving undead. But what about ant zombies? Yes, that's right: ant zombies.


Image: new bird discovered in Madagascar
(02/24/2011) The rich and unique biodiversity of Madagascar has a new member: a forest dwelling bird in the rail family, dubbed Mentocrex beankaensis. In 2009 US and Malaygasy scientists conducted a survey in Madagascar's dry Beanka Forest. They discovered several new species, of which the new rail is the first to be described.


Worldwide search for 'lost frogs' ends with 4% success, but some surprises
(02/16/2011) Last August, a group of conservation agencies launched the Search for Lost Frogs, which employed 126 researchers to scour 21 countries for 100 amphibian species, some of which have not been seen for decades. After five months, expeditions found 4 amphibians out of the 100 targets, highlighting the likelihood that most of the remaining species are in fact extinct; however the global expedition also uncovered some happy surprises. Amphibians have been devastated over the last few decades; highly sensitive to environmental impacts, species have been hard hit by deforestation, habitat loss, pollution, agricultural chemicals, overexploitation for food, climate change, and a devastating fungal disease, chytridiomycosis. Researchers say that in the past 30 years, its likely 120 amphibians have been lost forever.


Photos: new super tiny frogs discovered in Sri Lanka, one critically endangered
(02/09/2011) Two new incredibly small frogs have been discovered in Sri Lanka, an island nation off India with at least 100 species of frogs. The new frogs are in the genus Pseudophilautus, which are shrub frogs native either to Sri Lanka or India. One of the new species, dubbed Pseudophilautus hankeni, survives only in high mountain forests, and will likely be classified as Critically Endangered. "These species were discovered as a part of a broad amphibian survey that we carried out about 10 years ago in Sri Lanka. In that survey we discovered nearly 100 new species new to science. We are in the process of describing them now," explained Dr. Madhava Meegaskumbura, who participated in the frogs' discovery and formal description, to mongabay.com.


Egyptian jackal is actually ancient wolf
(01/26/2011) The Egyptian jackal, which may have been the inspiration for the Egyptian god Anubis, is actually not a jackal at all but a member of the wolf family. New genetic research in the open-access journal PLoS ONE finds that the Egyptian jackal is Africa's only member of the gray wolf family. The new wolf, dubbed by researchers as the African wolf, is most closely related to the Himalayan wolf.


Picture: scientists identify first known single-fingered dinosaur
(01/25/2011) Paleontologists working in China have discovered a first for dinosaurs: a species with only one finger. Named Linhenykus monodactylus, the extinct species stood only about two feet high and weighed about as much as a large parrot. Although small, the new dinosaur was a member of the carnivorous therapod dinosaurs, which include the infamous Tyrannosaurus Rex. The find was announced in the Proceedings of the National Academy of Sciences.


Africa gains new elephant species
(01/19/2011) DNA evidence has shown that the forest elephant-Africa's smaller, shyer pachyderm-is indeed a separate species from the much more well-known savanna elephant. While scientists have long debated the status of the forest elephant (should it be considered a separate population, a subspecies, or a unique species?) a new study in the open-access journal PLoS Biology finds that genetically the forest elephant is unarguably a new species. If conservation authorities accept the new study, it will change elephant conservation efforts throughout Africa.


Top 10 Environmental Stories of 2010
(12/20/2010) Below is a quick review of some of the biggest environmental stories of 2010: Climate change rears it ugly head; Oil spill in the Gulf; Agreement to save global biodiversity; Illegal logging crisis in Madagascar; REDD kicks off in Indonesia; Brazil deforestation falls to its lowest level; Hungary's red sludge; Nestle caves to social media activists; New mammals galore' and Global climate framework back on the table?


Picture: New lemur in Madagascar
(12/15/2010) Researchers have discovered a new species of lemur in Madagascar.


Genetic analysis uncovers new parrot on the edge of extinction
(11/23/2010) Down to just over 100 individuals, DNA analysis has revealed one of the world's most imperiled bird species: the western ground parrot Pezoporus flaviventris.. Genetic evidence collected from museums specimens, some well-over a century old, have led scientists to "cautiously suggest" that Australia's ground parrot be split into two distinct species—the eastern and the western—and not subspecies as they are currently considered. According to the study, the ground parrot species split apart some two million years ago, around the same time as the first members of the genus Homo evolved.


Photo: expedition discovers 20% of world's squids, including new species
(11/18/2010) An expedition to the seamounts of the southern Indian Ocean has proven that the region is a biodiverse hotspot for squids. To date, the expedition has identified 70 species of squid comprising 20% of the world's known squid species. But that's not all: they have also uncovered new species. At just over 2 feet long (27 inches or 70 centimeters), a species of squid found by the expedition proves to a brand new member of the chiroteuthid family. Squids from this family, which number around a dozen known species, employ bioluminescent organs to attract unwary prey.


New bat species confirmed in Ecuador, may already be extinct
(11/16/2010) Although the first specimen was collected over 30 years ago, scientists have only now confirmed that a tiny brown bat is indeed a unique species. Named Myotis diminutus for its incredibly small size, the new bat was discovered in the Chocó biodiversity hotspot, amid the moist forests of western Ecuador.


Photographer discovers new species of meat-eating plant in Cambodia
(11/16/2010) British photographer Jeremy Holdren recently discovered a new species of carnivorous pitcher plant in Cambodia's Cardamom Mountains during a survey with Fauna & Flora International (FFI).


Pictures: 'Mr. Burns' frog discovered in Colombia along with 2 other new species
(11/15/2010) Three previously undocumented species of frog have been discovered in Colombia, reports Conservation International.


Chaco biodiversity expedition suspended
(11/15/2010) A joint expedition by the Natural History Museum (NHM), London and the Natural History Museum, Asuncion to the dwindling dry forest of the Gran Chaco in Paraguay to record biodiversity, and hopefully uncover 'hundreds' of new species, has been suspended by the Paraguayan government. The suspension comes after a local organization voiced concern that the expedition would threaten uncontacted member of the Ayoreo tribe in the forest.


Chaco expedition working to "minimize the risk" of running into uncontacted natives
(11/11/2010) A joint expedition by the Natural History Museum (NHM), London and the Natural History Museum, Asuncion to study the biodiversity of the dwindling dry forests of Chaco in Paraguay have responded to recent concerns that they risk encountering uncontacted natives, which could potentially threaten the natives' lives as well as their own.


Eight new plants discovered in Bolivia
(11/07/2010) Researchers have described eight new species of plant from in and near Madidi National Park in the Bolivian Andes. Described in the journal Novon by botanists with the Missouri Botanical Garden and the National Herbaium in Laz Paz, Bolivia, seven of the eight plants were found as apart of the Proyecto Madidi (Project Madidi), a ten year effort to describe the plant species of three inter-connecting protected areas in Bolivia—Madidi National Park, Pilón Lajas Biosphere Reserve and Communal Lands, and Apolobamba Integrated Management Natural Area.


Picture: new monkey discovered in Myanmar
(10/26/2010) Hunters' reports have led scientists to discover a new species of monkey in the northern forests of Myanmar. Discovered by biologists from the Myanmar Biodiversity and Nature Conservation Association with support from primatologists with Fauna & Flora International (FFI) and the People Resources and Biodiversity Foundation, the strange looking primate is a member of the snub-nosed monkey family, adding a fifth member to this unmistakably odd-looking group of Asian primates. However, the species survives in only a small single population, threatened by Chinese logging and hunting.


Life shocker: new species discovered every three days in the Amazon
(10/26/2010) A new report by the World Wide Fund for Nature (WWF) confirms the Amazon rainforest, even as it is shrinking due to deforestation, remains among the world's most surprising places. According to the report, Amazon Alive, over the past decade (1999-2009) researchers have found 1,200 new species in the Amazon: one new species for every three days. Not surprisingly invertebrates, including insects, made up the bulk of new discoveries. But no type of species was left out: from 1999-2009 researchers discovered 637 new plants, 357 fish, 216 amphibians, 55 reptiles, 39 mammals, and 16 new birds. In new discoveries over the past decade, the Amazon has beaten out a number of high-biodiversity contenders including Borneo, the Eastern Himalayas, and the Congo rainforest.


Photos: three new forest frogs discovered in Tanzania
(10/19/2010) Africa's most biodiverse nation, Tanzania, has added a few more species to its dockets. Researchers have discovered three new amphibians in the always surprising Eastern Arc Mountains, a region which has supplied a number of new species recently. All three new species are members of the frog genus Callulina. Described in the Zoological Journal of the Linnean Society the researchers warn that all three of the new frogs are confined to small habitats threatened by deforestation and firewood collection.


Photos: weird new species discovered in deep sea trench
(10/18/2010) Fish were not expected to be able to survive so deep, but scientists have captured footage of a new species of a scavenger-hunting snailfish swimming at an astounding 7,000 meters below the surface. The video, taken from an 8,000 meter-deep sea trench in the Southeast Pacific Ocean, showed a level of biodiversity that surprised seasoned marine biologist, who have previously surveyed five other deep sea trenches.


Video: New carnivorous swamp beast discovered in Madagascar
(10/15/2010) Mongabay.com's Rhett Butler provides an update on the forest news for the week ending October 15, 2010.


Picture: unknown carnivore discovered in Madagascar lake
(10/10/2010) Researchers have identified a previously unknown species of carnivore lurking in one of the world's most endangered lakes. Durrell's vontsira (Salanoia durrelli), named in honor of the late conservationist and writer Gerald Durrell, was first photographed swimming in Madagascar's Lake Alaotra in 2004. Subsequent surveys by scientists at the the Durrell Wildlife Conservation Trust, the Natural History Museum, London, Nature Heritage, Jersey, and Conservation International confirmed the mongoose-like creature was indeed a new species.


Photos: 200 new species discovered in 60-day expedition in New Guinea
(10/06/2010) A 2009 expedition to Papua New Guinea proves once again that the island-nation is as diverse in life as it is in human cultures. It took researchers with Conservation International (CI) and the local Institute for Biological Research (IBR) just two months to uncover a startling 200 new species: averaging more than 3 a day in the remote Nakanai Mountains and Muller Range rising from the island of New Britain, a part of Papua New Guinea. Half of the new species were spiders, but the team also found two new mammals, nine new plants, two dozen frogs, and multitude of insects. Most surprising was the discovery of at least two species so unique that they are likely to be assigned their own genus.


Number of described marine species climbs from 230,000 to 250,000
(10/05/2010) After a decade of research in oceans around the world, scientists released the final results of the first Census of Marine Life (CoML) on Monday. The research, which involved 2,700 scientists from more than 80 countries, increased the number of known marine species from 230,000 to 250,000.


New ape species uncovered in Asia
(09/21/2010) Discovering a species unknown to science is a highlight of any biologist's career, but imagine discovering a new ape? Researchers with the German Primate Center (DPZ) announced today the discovery of a new species of ape in the gibbon family, dubbed the northern buffed-cheeked gibbon (Nomascus annamensis), according to the AFP. The new species was discovered in rainforests between the borders of Vietnam, Laos, and Cambodia: an area that contains a number of gibbon species.


Photos: New species spins the world's strongest and largest spider web
(09/20/2010) A newly discovered species of spider in Madagascar spins the world's longest web, report scientists writing in the Journal of Arachnology and PLoS ONE.


How the overlooked peccary engineers the Amazon, an interview with Harald Beck
(09/20/2010) When people think of the Amazon rainforest, they likely think of roaring jaguars, jumping monkeys, marching ants, and squeezing anacondas. The humble peccary would hardly be among the first animals to cross their mind, if they even know such pig-like animals exists! Yet new research on the peccary is proving just how vital these species are to the world's greatest rainforest. As seed dispersers and seed destroyers, engineers of freshwater habitats and forest gaps, peccaries play an immense, long overlooked, role in the rainforest. "Peccaries have the highest density and biomass of any Neotropical mammal species. Obviously these fellows have quite an appetite for almost anything, but primarily they consume fruits and seeds. Their specialized jaws allow them to crush very hard seeds. The cracking sounds can be heard through the thick vegetation long before we could see them. As peccary herds bulldoze through the leaf litter in search for insects, frogs, seeds, and fruits, they destroy (i.e. snap and trample) many seedlings and saplings, sometimes leaving only the bare ground behind," Harald Beck, assistant professor at Towson University in Maryland, told mongabay.com in an interview.


Unknown elephant relative photographed in Kenya
(09/16/2010) Scientists conducting research in the Boni-Dodori forest on the coast of northeastern Kenya may have discovered a new species of giant elephant-shrew, reports the Zoological Society of London (ZSL).


Photos: 'Tarzan' chameleon discovered in Madagascar
(08/31/2010) Scientists have discovered a new species of chameleon in a small block of rainforest in Madagascar.


Photos: Asia's tiniest frog discovered living inside carnivorous plants in Borneo
(08/25/2010) One of the world's smallest frogs has been discovered living inside carnivorous plants in Borneo, reports Conservation International, a conservation group that is jointly supporting a campaign with IUCN to search for some of the world's 'lost amphibians.' The species, described in Zootaxa by Indraneil Das and Alexander Haas of the Institute of Biodiversity and Environmental Conservation at the Universiti Malaysia Sarawak and Biozentrum Grindel und Zoologisches Museum of Hamburg, is named Microhyla nepenthicola after the plant in which is was found, Nepenthes ampullaria, a species of pitcher plant from Malaysian Borneo.


Exploring Kenya's sky island
(08/18/2010) Rising over 2,500 meters from Kenya's northern desert, the Mathews Range is a sky island: isolated mountain forests surrounded by valleys. Long cut off from other forests, 'sky islands' such as this often contain unique species and ecosystems. Supported by the Nature Conservancy, an expedition including local community programs Northern Rangelands Trust and Namunyak Conservancy recently spent a week surveying the mountain range, expanding the range of a number of species and discovering what is likely a new insect.


Stunning monkey discovered in the Colombian Amazon
(08/11/2010) While the Amazon is being whittled away on all sides by logging, agriculture, roads, cattle ranching, mining, oil and gas exploration, today's announcement of a new monkey species proves that the world's greatest tropical rainforest still has many surprises to reveal. Scientists with the National University of Colombia and support from Conservation International (CI) have announced the discovery of a new monkey in the journal Primate Conservation on the Colombian border with Peru and Ecuador. The new species is a titi monkey, dubbed the Caquetá titi ( Callicebus caquetensis). However, the announcement comes with deep concern as researchers say it is likely the new species is already Critically Endangered due to a small population living in an area undergoing rapid deforestation for agriculture.


What is the most unexplored ecosystem on Earth?
(08/02/2010) One should be forgiven for thinking that the most under-explored ecosystem in the world is tropical rainforests or cave systems since new species are frequently uncovered in such places. But new research in the open-access journal PLoS ONE finds that the least explored part of the world is the deep sea, especially what lies beneath the open ocean. Ironically, not only does this ecosystem remain the least explored, it is also the world's largest habitat.


Eleven new species discovered in France
(07/29/2010) Usually announcements of new species come from biodiverse rainforests or unexplored marine depths, but researchers have announced the discovery of nearly a dozen new species in one of Earth's most well-trodden place: France. Eleven new species have been discovered in Mercantour National Park in southern France. All the new species are insects, including one beetle, seven new aquatic invertebrate living under creek beds, and three springtails, which are soil-dwelling arthropods.


Two new frogs discovered in Panama amidst amphibian plague
(06/06/2010) Researchers working to save Panama's frogs from a fatal disease have stumbled on two species unknown to science. In Omar Torrijos National Park they found a bigger version of a common species, which is now known to be a unique species, and near the Colombian border they discovered a new frog that has been named after Spanish for DNA. Both frogs were discovered while researchers searched for frog populations in chytridiomycosis-infected areas. The highly contagious disease chytridiomycosis has devastated frog species worldwide and is believe to be at least in part responsible for some 100 extinctions of amphibians.


18,225 new species discovered in 2008
(05/27/2010) In the 2010 State of Observed Species researchers have announced that 18,225 living species were discovered in 2008. In addition, 2,140 new extinct species were discovered byway of fossils.


New bird discovered in Colombia—and released alive
(05/26/2010) Researchers have discovered a new species of antpitta in the montane cloud forests of the Colibri del Sol Bird Reserve in western Colombia. A thrush-like bird, the new cinnamon and gray species was, according to a press release by the American Bird Conservancy (ABC), "captured, banded, measured, photographed, sampled for DNA, and then released alive back into the wild". This is one of only a few incidences in which a new species has been described without 'collecting' an individual (i.e. killing) to provide a model of the species in a museum. The new bird has been named Fenwick's antpitta (Grallaria fenwickorum) after the President of ABC, George Fenwick, and his family.


Photos: the penis-like mushroom and other top 10 new species of 2009
(05/23/2010) The International Institute for Species Exploration at Arizona State University has released its annual top 10 list of new species discovered last year. This time the list includes a two inch penis-like mushroom, a minnow named after Bram Stoker's world-famous horror-character, a bomb-throwing deep sea worm, a giant carnivorous plant named after TV personality and conservationist David Attenborough, and a beautifully patterned frogfish.













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